A Cellular U-Turn: How a Social Bacterium Hits Reset After Division

Discover how Myxococcus xanthus performs a complete "factory reset" on its internal navigation system after every cell division.

Biology Research 8 min read October 2023

Introduction: The Microscopic World is Not a Random Mess

Imagine a city where every time a new building was completed, all the doors, roads, and traffic signals reset to a blank slate. Chaos would ensue, but only temporarily, before a new, organized layout emerged. In the microscopic world, something remarkably similar happens.

For cells, "front" and "back" aren't arbitrary—this polarity is crucial for movement, feeding, and communication. For the social bacterium Myxococcus xanthus, a master of coordination, this process is vital. Recent research reveals a fascinating secret: every time it divides, it performs a complete "factory reset" on its internal navigation system. This isn't a bug; it's a brilliant feature that ensures its survival.

Key Insight

Cell polarity defines directionality in cellular processes, much like a compass guides navigation.

The Reset

Division erases all existing polarity, allowing each new cell to establish its own direction independently.

The Social Life of a Gliding Bacterium

Before we dive into the reset, let's meet our protagonist. Myxococcus xanthus is a soil-dwelling bacterium, often called a "social bacterium" or "wolf pack" hunter because it swarms collectively to prey on other microbes.

How Does It Move?

Unlike many bacteria that swim with whip-like flagella, M. xanthus glides smoothly over surfaces using two ingenious molecular engines:

Adventurous (A-Motility) System

This engine is like a tank tread. Imagine molecular treads inside the cell's membrane that circulate, gripping the surface and propelling the bacterium forward. This system allows individual cells to strike out on their own.

Social (S-Motility) System

This is more like a grappling hook. The bacterium extends a sticky, spear-like pilus from its front, attaches to a surface or another cell, and then retracts it to pull itself along. This is essential for moving in coordinated groups.

Both of these systems require the cell to have a defined front (leading edge) and back (lagging edge). This polarity is established by a protein called MglA, a molecular switch that, when active (bound to GTP), localizes to the front of the cell and tells the motility machinery where to assemble.

The Puzzling Question: What Happens at Division?

When a rod-shaped bacterium like M. xanthus prepares to divide, it elongates and then pinches in the middle to create two daughter cells. This raised a critical question for scientists: At the moment of division, each daughter cell inherits one "old" pole (from the mother cell) and one "new" pole (the division site). Does the existing polarity from the mother cell get passed down, or is it erased and re-established from scratch?

In-Depth Look: The Key Experiment That Captured the Reset

A crucial experiment by scientists at the University of Basel provided a clear answer. They set out to watch polarity in real-time throughout the entire cell division process.

Methodology: Lighting Up the "Steering Wheel"

The researchers used a powerful combination of tools to make the invisible visible.

Fluorescent Tagging

They genetically engineered M. xanthus cells so that the key polarity protein, MglA, was fused to a green fluorescent protein (GFP). When placed under a high-resolution fluorescence microscope, the MglA protein would glow green, revealing its precise location inside the cell.

Time-Lapse Microscopy

They placed the glowing bacteria on a nutrient-poor surface and filmed them for several hours under the microscope, capturing an image every few minutes.

Cell Tracking

They carefully tracked individual cells as they moved, grew, and eventually divided. By analyzing the videos, they could see exactly where the bright green spot of MglA was located before, during, and after division.

Research Toolkit
Tool / Reagent Function in the Experiment
Fluorescent Protein (GFP) A molecular "tag" that is fused to a protein of interest (like MglA), allowing scientists to visualize its location and movement inside a living cell under a microscope.
Time-Lapse Fluorescence Microscopy A technique for taking repeated images of living cells over time. It was essential for capturing the dynamic process of polarity loss and regain during division.
MglA Mutants Genetically altered versions of the MglA protein that are "stuck" in an always-on or always-off state. These were used to confirm MglA's role as the master polarity switch.
Minimal Agar Plates A simple, nutrient-poor gel surface on which the bacteria are grown for microscopy. It encourages motility and ensures the cells remain flat and in focus for imaging.

Results and Analysis: A Clear Pattern Emerges

The results were striking and consistent. The data told a clear story of erasure and rebirth.

  • Before Division: A motile mother cell had a single, bright focus of MglA-GFP at one pole, defining it as the "front."
  • During Division: As the cell constricted in the middle, the MglA signal at the old pole disappeared. For a brief period, the dividing cell had no visible polarity—it was, in effect, unpolarized and non-motile.
  • After Division: Immediately following separation, each newborn daughter cell independently established a new MglA focus. Critically, this new "front" could be at either the old pole or the new pole, with a near-perfect 50/50 probability.

This was the smoking gun. If polarity were simply inherited, one daughter would always keep the old front, and the other would be born with a random front. The complete erasure and random re-establishment proved that cell division acts as a hard reset for cellular polarity.

MglA Polarity During Cell Division
Cell Stage MglA Localization Cell Motility
Mother Cell Single focus at one pole Motile
Dividing Cell Signal lost; no focus Non-motile
Daughter Cells New focus at random pole Motile
Front Selection in Daughter Cells
Consequences of Disrupting the Reset Mechanism
Experimental Condition Effect on Polarity Reset Outcome for Population
Normal Wild-Type Cells Clean reset after division Healthy mix of movement directions; efficient swarming
Mutant Cells (defective reset) Polarity inherited or stuck Daughters move in same direction; chaotic, inefficient movement; failed swarm expansion

Conclusion: Why Hit Reset? The Bigger Picture

This reset is more than just a curious cellular quirk; it's a profound survival strategy. By randomizing direction after every division, the bacterial population ensures maximum exploration of its environment. If both daughters always followed the mother's path, the swarm would be a narrow, single-file line, easily missing nearby prey.

Instead, the reset creates a constantly expanding, dynamic front, perfect for hunting as a pack.

The discovery that a fundamental process like cell division includes a step to erase cellular memory teaches us a broader lesson in biology: Order often emerges from a deliberate step back into chaos. For Myxococcus xanthus, this cyclical dance of structure and reset is the key to its successful, social life.

The Cellular Reset Principle

The deliberate erasure of polarity during division ensures population diversity in movement direction, enhancing environmental exploration and collective survival.

Key Findings
  • Cell division resets cellular polarity completely
  • MglA protein localization is erased during division
  • New polarity is established randomly in daughter cells
  • This reset enhances swarm exploration efficiency
Polarity Reset Process
Related Concepts
Cell Polarity Bacterial Motility Protein Localization Fluorescence Microscopy Collective Behavior MglA GTPase